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Russian missiles 'fell on Iran' Syria crisis: Russian missiles 'fell on Iran'
(35 minutes later)
Four Russian cruise missiles fired at Syria from the Caspian Sea landed on Iran, unnamed US officials say Four Russian cruise missiles fired at Syria from the Caspian Sea landed in Iran, unnamed US officials say.
This breaking news story is being updated and more details will be published shortly. Please refresh the page for the fullest version. It was unclear whether the missiles caused any damage, they said.
If you want to receive Breaking News alerts via email, or on a smartphone or tablet via the BBC News App then details on how to do so are available on this help page. You can also follow @BBCBreaking on Twitter to get the latest alerts. Russia's defence ministry has declined to comment. On Wednesday, Russia said it had launched 26 cruise missiles at targets in north and north-west Syria.
The news came as Nato renewed assurances to defend its allies in view of the "escalation of Russian military activities" in Syria.
Nato is boosting its response forces to be able to deploy troops speedily.
Moscow denies Western accusations that it has mainly targeted opponents of Syrian President Bashar al-Assad, insisting its strikes have hit the infrastructure of the so-called Islamic State (IS) and other militant groups.
IS militants have seized swathes of territory in Syria and Iraq.
The Russian air strikes had "weakened" IS, Syrian Army Chief of Staff Gen Ali Abdullah Ayoub said on Thursday, enabling the army to start a "big attack" to retake towns and villages.
Heavy fighting was reported in areas of Idlib, Hama and Latakia provinces, where a coalition of rebels that includes the Nusra Front operates.
Government-backed troops had moved into the key Ghab plain area, the UK-based Syrian Observatory for Human Rights monitoring group said.